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Department of Paleobiology

  • Scott L. Wing
  • Curator of Fossil Plants
  • Phone:   202-633-1361
  • Fax:   202-786-2832
  • E-mail Address:   wings
  • Mailing Address:
    Smithsonian Institution
    PO Box 37012, MRC 121
    Washington, DC 20013-7012
  • Shipping Address:
    Smithsonian Institution
    National Museum of Natural History
    10th & Constitution NW
    Washington, DC 20560-0121


Ph.D. Yale University, 1981
B.S. Yale College, 1976

Research Interests

Descriptions and photos of my research projects (link).

I am a biologist interested in evolution and ecology. I study fossils because they provide a long-term record of evolutionary, ecological and environmental change. If properly interpreted that record can say much about process as well as history. I use fossil plants to reconstruct past climates and local environments, and I also study the composition and diversity of floras through time as they respond to changing conditions. My research has focused on the Cretaceous and early Cenozoic - a period of globally warm climate when flowering plants were emerging as the dominant form of terrestrial life. My research is based largely on field work and collections made by me or under my direction. I collect data on the morphology and taxonomy of fossil and analyze them through statistical characterization of trends in morphology, composition or diversity of floras, and comparison of floral change with indicators of environmental change. I also make climatic interpretations based on fossil plants for comparison with paleoclimatic estimates derived from computer simulations.

I have a long-running project examining climatic and floral change across the Paleocene/Eocene boundary, a time of global climatic warming. The project has been directed at quantifying temperature and precipitation change and also change in floral composition and diversity over about 5 million years spanning the boundary. Much of my field work has been in western North America because there are abundant fossils, the stratigraphic and temporal context of the fossils has been established (by me and many others), and study of these areas is interdisciplinary. I have also worked on this project in Pakistan and Argentina.

In a second long-term project I am studying the abundance and diversity of angiosperms in the Late Cretaceous. Angiosperms are the youngest major group of terrestrial organisms, which raises the question: how did they achieve such high diversity and abundance over a relatively brief geological interval (~100 - 60 million years ago)? I have been examining exceptionally preserved, in situ, fossil floras from the Cretaceous in order to establish the relative contribution of angiosperms to vegetational biomass and overall plant diversity in different types of environments. I am using these data to understand the ecological dimensions of this major evolutionary radiation.

I am also a member of a large team of paleobotanists who are adding data (mostly from the literature) on fossil plant assemblages to a web-accessible database maintained at the National Center for Ecological Analysis and Synthesis. In the long run this database will allow us to better quantify changes in the diversity and composition of terrestrial floras across the whole history of life on land. By comparing results compiled from the literature with more detailed studies like the ones I am doing in the Cretaceous and Cenozoic, I hope we will also reach a better understanding of the biases and limitations of the published fossil record.

Why is this research important?
Like any scientific research, my work is important to different constituencies for different reasons.
Humans are adding CO2 and other greenhouse gases to the atmosphere, which will change global climate. Climatologists estimate the rate and magnitude of future climate change using general circulation models (GCMs). How accurate are GCMs? Strictly, their predictions can only be tested by waiting to see how climate changes in the decades and centuries ahead, but some idea of their accuracy can be gained by comparing "postdictions" for past times against climate reconstructions for those past times based on fossil plants. Thus my paleoclimate reconstructions based on fossil plants are important to climatologists, and, to the extent that they reveal strengths and weaknesses in GCMs, to policy makers and the public as well.

I have an academic interest in floral change during past periods of global climate, but this work may have practical significance if it reveals the rates and mechanisms by which plants respond to climate change, since these processes are important in predicting the effect of future anthropogenic climate change on natural populations.

Finally, I think there is intrinsic importance to learning about the history of the earth and the organisms that inhabit it. This knowledge can bring about in all of us a greater appreciation for the complexity of nature, it gives us context for our own existence, and it reveals the uniqueness of our current situation - in which we hold a degree of stewardship over the planet.

Research Assistants, Current and Former Post-doctoral Fellows, and Students

Post-Doctoral Fellows:

Graduate Students and Pre-Doctoral Fellows

  • 2009- Andy Simpson, Behavior, Ecology, Evolution & Systematics Program, University of Maryland College Park (coadvisor)
  • 2008- Nathan Jud, Behavior, Ecology, Evolution & Systematics Program, University of Maryland College Park (Peter Buck Pre-doctoral Fellow)
  • 2003-2008 Ellen Currano, Department of Geosciences, Pennsylvania State University (coadvisor)
  • 2000-2008 Mandela Lyon, Ph.D. candidate, Dept. of Earth and Environmental Science, University of Pennsylvania
  • 1994-1998 Peter Wilf, Department of Geology, University of Pennsylvania (primary thesis advisor)

Summer Interns

  • 2011 James Super, Stanford University
  • 2010 Leah Rand, University of Chicago, Featured in the film below!
  • 2009 Ben Casterline and Eric Mayer, University of Chicago
  • 2008 Marie Hoerner and Hayden Miller, Colorado College; James Super, Stanford University
  • 2007 James Super, Stanford University
  • 2006 Shannon Loomis, University of Chicago
  • 2005 Elizabeth Lovelock, University of Pennsylvania
  • 2004 Anthony Alvarez, University of California Santa Barbara
  • 2002 Ellen Currano, University of Chicago
  • 2001 David Blackburn, University of Chicago
  • 2000 Courtenay Shinn, University of Chicago
  • 1999 Noel Heim, University of Chicago
  • 1999 Jessie Scott, Wellesley College
  • 1998 Josh Chamot, College of William & Mary
  • 1988-1996 (23 students from various colleges)


  • Adjunct Professor, Department of Biology and BEES (Behavior, Ecology, Evolution, and Systematics) Group, University of Maryland College Park
  • Courtesy Curator, Florida Museum of Natural History, University of Florida, Gainseville
  • Honorary Senior Research Fellow, Geography and Environmental Sciences, University of Birmingham
  • Adjunct Professor, Department of Geological Sciences, University of Michigan
  • Department of Earth and Environmental Sciences and Geobiology Group, University of Pennsylvania

Volunteer Positions

  • 2006-2008 Editorial Board, Palaios
  • 2006-2008 Editorial Board, Palaeoworld
  • 2001-2004 Editorial Board, Evolutionary Ecology Research
  • 1999-2003 Editorial Board, Annual Reviews of Ecology and Systematics
  • 1995-2001 Co-Editor, Paleobiology, and member of Paleontological Society Council
  • 1996-1997 President, Paleontological Society of Washington, D.C.
  • 1996-1997 Chair, NMNH Public Programs Committee
  • 1995-1997 Paleontological Society Committee for the Geobiology of Critical Intervals
  • 1994-1996 Organizing and Program Committees, North American Paleontological Convention VI
  • 1994-1995 Chairman, Senate of Scientists, National Museum of Natural History
  • 1994 Secretary, Smithsonian Institution Congress of Scholars
  • 1994-1995 Environmental Geoscience Advisory Committee, American Geological Institute
  • 1993-1994 Treasurer, Senate of Scientists, National Museum of Natural History
  • 1990-1993 Co-director, National Museum of Natural History Research Training Program

Symposia/Meetings Organized

  • Climate and Biota of the Early Paleogene. June 2006. Scientific Committee Member. Bilbao, Spain.
  • Causes and Effects of the Paleocene-Eocene Thermal Maximum and Other Paleogene Hyperthermal Events. October 2005. Organized by Scott Wing. Geological Society of America Annual Meeting, Salt Lake City
  • A Global View of Paleogene Floras. March 2004. Organized by Peter Wilf, Kirk Johnson, and Scott Wing. Seventh International Organization of Paleobotany Conference, Bariloche, Argentina.
  • Disruptions in Ancient Land Ecosystems: Lessons from the Fossil Record. February 2003. Organized by A.K. Behrensmeyer and S. L. Wing. AAAS Annual Meeting, Denver, Colorado.
  • Climate and Biota of the Early Paleogene. July 2001. Organized by Scott Wing, Philip Gingerich & Amanda Ash. Sponsored by NSF and National Museum of Natural History, Smithsonian Institution.
  • Globally Warm Climates of the Early Cenozoic: Evidence, Causes and Biotic Consequences. October 1999. Organized by Scott Wing and Lisa Sloan. Geological Society of America Annual Meeting, Pardee Symposium.
  • Fifth International Organization of Paleobotany Meeting. Vegetation, Climate, and Phytogeography during the Eocene "Greenhouse" Interval. June 1996. Organized by Scott Wing and R. J. Burnham.
  • North American Paleontological Convention VI. Model Results and Paleontological Inferences for Greenhouse Intervals in Earth History. June 1996. Organized by Brian Huber, Ken MacLeod & Scott Wing.
  • Long-term and short-term views of community stability: implications for evolution. July 1990. Organized by W. A. DiMichele and Scott Wing. 4th Int. Congress of Systematics and Evolutionary Biology.
  • Global Change in Earth History, March 1990. National Museum of Natural History Briefing for Congress and Federal Agencies. Organized by Scott Wing. Sponsored by the Smithsonian Office of External Affairs.
  • First International Symposium on the Evolution of Terrestrial Ecosystems - The Current Status of Terrestrial Paleoecology. May 1987. Organized by A. K. Behrensmeyer, Scott Wing, W.A. DiMichele and R. Potts. Sponsored by the Evolution of Terrestrial Ecosystems Program, Smithsonian Institution.
  • Short Course on Paleoecological Methods. August 1987. Organized by W.A. DiMichele and Scott Wing. Sponsored by the Paleobotanical Section of the Botanical Society of America.
  • Conference and workshop on peat forming environments. January 1987. Organized by Scott Wing and W.A. DiMichele. Cosponsored by the Evolution of Terrestrial Ecosystems Program and Utah International.

[ TOP ]


Barclay, Richard S. and Wing, Scott L. 2016. Improving the Ginkgo CO2 barometer: Implications for the early Cenozoic atmosphere, Earth and Planetary Science Letters, 439:158-171

Wilf, Peter, Zhang, Shengping, Chikkerur, Sharat, Little, Stefan A., Wing, Scott L. and Serre, Thomas. 2016. Computer vision cracks the leaf code, Proceedings of the National Academy of Sciences, 113(12):3305-3310

Zalasiewicz, Jan, Waters, Colin N., Williams, Mark, Barnosky, Anthony D., Cearreta, Alejandro, Crutzen, Paul, Ellis, Erle, Ellis, Michael A., Fairchild, Ian J., Grinevald, Jacques, Haff, Peter K., Hajdas, Irka, Leinfelder, Reinhold, McNeill, John, Odada, Eric O., Poirier, Clément, Richter, Daniel, Steffen, Will, Summerhayes, Colin, Syvitski, James P. M., Vidas, Davor, Wagreich, Michael, Wing, Scott L., Wolfe, Alexander P., An, Zhisheng, et al. 2015. When did the Anthropocene begin? A mid-twentieth century boundary level is stratigraphically optimal, Quaternary International, 283:196-203

Bowen, Gabriel J., Maibauer, Bianca J., Kraus, Mary J., Rohl, Ursula, Westerhold, Thomas, Steimke, Amy, Gingerich, Philip D., Wing, Scott L. and Clyde, William C. 2015. Two massive, rapid releases of carbon during the onset of the Palaeocene-Eocene thermal maximum, Nature Geoscience, (8):44-47

Bocxlaer, Bert V., Clewing, Catharina, Etimosundja, Jean-Papy Mongindo, Kankonda, Alidor, Ndeo, Oscar Wembo and Albrecht, Christian. 2015. Recurrent camouflaged invasions and dispersal of an Asian freshwater gastropod in tropical Africa, BMC Evolutionary Biology, 15

Diefendorf, Aaron F., Freeman, Katherine H., Wing, Scott L., Currano, Ellen D. and Mueller, Kevin E. 2015. Paleogene plants fractionated carbon isotopes similar to modern plants, Earth and Planetary Science Letters, 429:33-44

Lyons, Sara K., Amatangelo, Kathryn L., Behrensmeyer, Anna K., Bercovici, Antoine, Blois, Jessica L., Davis, Matt, DiMichele, William A., Du, Andrew, Eronen, Jussi T., Faith, J. Tyler, Graves, Gary R., Jud, Nathan, Labandeira, Conrad C., Looy, Cindy V., McGill, Brian, Miller, Joshua H., Patterson, David, Pineda-Munoz, Silvia, Potts, Richard, Riddle, Brett, Terry, Rebecca, Tóth, Anikó, Ulrich, Werner, Villaseñor, Amelia, Wing, Scott L., et al. 2015. Holocene shifts in the assembly of plant and animal communities implicate human impacts, Nature, (529):80-83

Diefendorf, Aaron F., Leslie, Andrew B. and Wing, Scott L. 2015. Leaf wax composition and carbon isotopes vary among major conifer groups, Geochimica et Cosmochimica Acta, 170:145-156

Green, Walton A., Little, Stefan A., Price, Charles A., Wing, Scott L., Smith, Selena Y., Kotrc, Benjamin and Doria, Gabriela. 2014. Reading the leaves: A comparison of leaf rank and automated areole measurement for quantifying aspects of leaf venation, Applications in Plant Sciences, 2(8)

Graham, Heather V., Patzkowsky, Mark E., Wing, Scott L., Parker, Geoffrey G., Fogel, Marilyn L. and Freeman, Katherine H. 2014. Isotopic characteristics of canopies in simulated leaf assemblages, Geochimica Et Cosmochimica Acta, 144:82-95

Blois, Jessica L., Gotelli, Nicholas J., Behrensmeyer, Anna K., Faith, J. T., Lyons, Sara K., Williams, John W., Amatangelo, Kathryn L., Bercovici, Antoine, Du, Andrew, Eronen, Jussi T., Graves, Gary R., Jud, Nathan, Labandeira, Conrad C., Looy, Cindy V., McGill, Brian, Patterson, David, Potts, Richard, Riddle, Brett, Terry, Rebecca, Tóth, Anikó, Villaseñor, Amelia and Wing, Scott L. 2014. A framework for evaluating the influence of climate, dispersal limitation, and biotic interactions using fossil pollen associations across the late Quaternary, Ecography, 37(11):1095-1108

Little, Stefan A., Green, Walton A., Wing, Scott L. and Wilf, Peter. 2014. Reinvestigation of Leaf Rank, an Underappreciated Component of Leo Hickey's Legacy, Bulletin of the Peabody Museum of Natural History, 55(2):79-87

Peppe, Daniel J., Lemons, Casee R., Royer, Dana L., Wing, Scott L., Wright, Ian J., Lusk, Christopher H. and Rhoden, Chazelle H. 2014. Biomechanical and leaf–climate relationships: A comparison of ferns and seed plants, American Journal of Botany, 101(2):338-347

Diefendorf, Aaron F., Freeman, Katherine H. and Wing, Scott L. 2014. A comparison of terpenoid and leaf fossil vegetation proxies in Paleocene and Eocene Bighorn Basin sediments, Organic Geochemistry, 71:30-42

Wing, Scott L., Johnson, Kirk R., Peppe, Daniel J., Green, Walton A. and Taylor, David Winship. 2014. The Multi-Stranded Career of Leo J. Hickey, Bulletin of the Peabody Museum of Natural History, 55(2):69-78

Herrera, Fabiany, Manchester, Steven R., Carvalho, Mónica, Jaramillo, Carlos A. and Wing, Scott L. 2014. Paleocene wind-dispersed fruits and seeds from Colombia and their implications for early Neotropical rainforests, Acta Palaeobotanica, 54(2):197-229

Kraus, Mary J., McInerney, Francesca A., Wing, Scott L., Secord, Ross, Baczynski, Allison A. and Bloch, Jonathan I. 2013. Paleohydrologic response to continental warming during the Paleocene-Eocene Thermal Maximum, Bighorn Basin, Wyoming, Palaeogeography, Palaeoclimatology, Palaeoecology, 370:196-208

Wing, Scott L. and Currano, Ellen D. 2013. Plant response to a global greenhouse event 56 million years ago, American Journal of Botany, 100(9):1234-1254

Clyde, W. C., Gingerich, P. D., Wing, Scott L., Röhl, U., Westerhold, T., Bowen, G., Johnson, K., Baczynski, A. A., Diefendorf, A., McInerney, F., Schnurrenberger, D., Noren, A. and Brady, K. 2013. Bighorn Basin Coring Project (BBCP): a continental perspective on early Paleogene hyperthermals, Scientific Drilling, 16:21-31

Baczynski, Allison A., McInerney, Francesca A., Wing, Scott L., Kraus, Mary J., Bloch, Jonathan I., Boyer, Doug M., Secord, Ross, Morse, Paul E. and Fricke, Henry C. 2013. Chemostratigraphic implications of spatial variation in the Paleocene-Eocene Thermal Maximum carbon isotope excursion, SE Bighorn Basin, Wyoming, Geochemistry, Geophysics, Geosystems, 14(10):4133-4152

Clyde, W. C., Gingerich, P. D. and Wing, Scott L. 2012. Coring Project in Bighorn Basin: Drilling Phase Complete, Eos, 93(4):41-42

Diefendorf, Aaron F., Freeman, Katherine H. and Wing, Scott L. 2012. Distribution and carbon isotope patterns of diterpenoids and triterpenoids in modern temperate C3 trees and their geochemical significance, Geochimica et Cosmochimica Acta, 85:342-356

Secord, Ross, Bloch, Jonathan I., Chester, Stephen G. B., Boyer, Doug M., Wood, Aaron R., Wing, Scott L., Kraus, Mary J., McInerney, Francesca A. and Krigbaum, John. 2012. Evolution of the Earliest Horses Driven by Climate Change in the Paleocene-Eocene Thermal Maximum, Science, 335(6071):959-962

Wing, Scott L., Stromberg, Caroline A. E., Hickey, Leo J., Tiver, Fleur, Willis, Brian, Burnham, Robyn J. and Behrensmeyer, Anna K. 2012. Floral and environmental gradients on a Late Cretaceous landscape, Ecological Monographs, 82(1):23-47

Price, Charles A., Wing, Scott L. and Weitz, Joshua S. 2012. Scaling and structure of dicotyledonous leaf venation networks, Ecology Letters, 15(2):87-95

Carvalho, Mónica R., Herrera, Fabiany, Jaramillo, Carlos A., Wing, Scott L. and Callejas, Ricardo. 2011. Paleocene Malvaceae from northern South America and their biogeographical implications, American Journal of Botany, 98(8):1337-1355

Wing, Scott L. 2011. Deep Future The Next 100,000 Years of Life on Earth, Science, 333(6044):825-825

McInerney, Francesca A. and Wing, Scott L. 2011. The Paleocene-Eocene Thermal Maximum: a Perturbation of Carbon Cycle, Climate, and Biosphere with Implications for the Future, Annual Review of Earth and Planetary Sciences, 39:489-516

Freeman, Katherine H., Mueller, Kevin E., Diefendorf, Aaron F., Wing, Scott L. and Koch, Paul L. 2011. Clarifying the influence of water availability and plant types on carbon isotope discrimination by C3 plants, Proceedings of the National Academy of Sciences of the United States of America, 108(16):E59-E60

Adams, Jason S., Kraus, Mary J. and Wing, Scott L. 2011. Evaluating the use of weathering indices for determining mean annual precipitation in the ancient stratigraphic record, Palaeogeography, Palaeoclimatology, Palaeoecology, 309(3-4):358-366

Diefendorf, Aaron F., Freeman, Katherine H., Wing, Scott L. and Graham, Heather V. 2011. Production of n-alkyl lipids in living plants and implications for the geologic past, Geochimica et Cosmochimica Acta, 75(23):7472-7485

Green, Walton A., Hunt, Gene, Wing, Scott L. and DiMichele, William A. 2011. Does extinction wield an axe or pruning shears? How interactions between phylogeny and ecology affect patterns of extinction, Paleobiology, 37(1):72-91

Wing, Scott L. 2011. We Need a Deeper Sense of Time, Science, 333(6044):825

Diefendorf, Aaron F., Mueller, Kevin E., Wing, Scott L., Koch, Paul L. and Freeman, Katherine H. 2010. Global patterns in leaf C-13 discrimination and implications for studies of past and future climate, Proceedings of the National Academy of Sciences of the United States of America, 107(13):5738-5743

Bennington, J. B., DiMichele, William A., Badgley, Catherine, Bambach, Richard K., Barrett, Paul M., Behrensmeyer, Anna K., Bobe, Rene, Burnham, Robyn J., Daeschler, Edward B., Van Dam, Jan, Eronen, Jussi T., Erwin, Douglas H., Finnegan, Seth, Holland, Steven M., Hunt, Gene, Jablonski, David, Jackson, Stephen T., Jacobs, Bonnie E., Kidwell, Susan M., Koch, Paul L., Kowalewski, Michal J., Labandeira, Conrad C., Looy, Cindy V., Lyons, Sara K., Novack-Gottshall, Philip M., et al. 2009. Critical Issues of Scale in Paleoecology, Palaios, 24(1-2):1-4

Wing, Scott L., Bloch, J. I., Bowen, G. J., Boyer, D. M., Chester, S., Diefendorf, Aaron F., Harrington, G. J., Kraus, M. J., Secord, R. and Smith, F. A. 2009. "Coordinated sedimentary and biotic change during the Paleocene-Eocene Thermal Maximum in the Bighorn Basin, Wyoming, USA". Pp. 156-162 in Climatic and Biotic Events of the Paleogene (CBEP 2009), extended abstracts from an international conference in Wellington, New Zealand, 12-15 January 2009 (Crouch, E. M., Strong, C. P. and Hollis, C. J.). Institute of Geological and Nuclear Sciences

Ellis, B., Daly, D. C., Hickey, Leo J., Johnson, Kirk R., Mitchell, J. D., Wilf, Peter and Wing, Scott L.2009. Manual of Leaf Architecture. Cornell University Press

Gomez-Navarro, Carolina, Jaramillo, Carlos A., Herrera, Fabiany, Wing, Scott L. and Callejas, Ricardo. 2009. Palms (Arecaceae) from a Paleocene rainforest of northern Colombia, American Journal of Botany, 96(7):1300-1312

Wing, Scott L., Herrera, Fabiany, Jaramillo, Carlos A., Gómez-Navarro, Carolina, Wilf, Peter and Labandeira, Conrad C. 2009. Late Paleocene fossils from the Cerrejón Formation, Colombia, are the earliest record of Neotropical rainforest, Proceedings of the National Academy of Sciences of the United States of America, 106(44):18627-18632

Currano, Ellen D., Wilf, Peter, Wing, Scott L., Labandeira, Conrad C., Lovelock, Elizabeth C. and Royer, Dana L. 2008. Sharply increased insect herbivory during the Paleocene-Eocene thermal maximum, Proceedings of the National Academy of Sciences of the United States of America, 105(6):1960-1964

Wing, Scott L., Bloch, Johathan I., Bowen, Gabriel J., Boyer, Douglas M., Chester, Stephen, Diefendorf, Aaron F., Harrington, Guy J., Kraus, Mary J., Secord, Ross and McInerney, Francesca A. 2008. "Coordinated Sedimentary and Biotic Change During the Paleocene-Eocene Thermal Maximum in the Bighorn Basin, Wyoming, USA". Pp. 157-163 in Climatic and Biotic Events of the Paleogene (CBEP 2009), extended abstracts from an international conference in Wellington, New Zealand, 12-15 January 2009 (Crouch, E. M., Strong, C. P. and Hollis, C. J.). Institute of Geological and Nuclear Sciences Limited

Herrera, Fabiany A., Jaramillo, Carlos A., Dilcher, David L., Wing, Scott L. and Gomez-N., Carolina. 2008. Fossil Araceae from a Paleocene Neotropical rainforest in Colombia, American Journal of Botany, 95(12):1569-1583

Secord, Ross, Wing, Scott L. and Chew, Amy. 2008. Stable isotopes in early Eocene mammals as indicators of forest canopy structure and resource partitioning, Paleobiology, 34(2):282-300

Clyde, William C., Hamzi, Walid, Finarelli, John A., Wing, Scott L., Schankler, David and Chew, Amy. 2007. Basin-wide magnetostratigraphic framework for the Bighorn Basin, Wyoming, Geological Society of America Bulletin, 119(7/8):848-859

Smith, Francesca A., Wing, Scott L. and Freeman, Katherine H. 2007. Magnitude of the carbon isotope excursion at the Paleocene-Eocene thermal maximum: The role of plant community change, Earth and Planetary Science Letters, 262(1-2):50-65

Bowen, Gabriel J., Bralower, Timothy J., Delaney, Margaret L., Dickens, Gerald R., Kelly, Daniel C., Koch, Paul L., Kump, Lee R., Meng, Jin, Sloan, Lisa C., Thomas, Ellen, Wing, Scott L. and Zachos, James C. 2006. Eocene Hyperthermal Event Offers Insight Into Greenhouse Warming, Eos, 87(17):165-167

McMillan, M. E., Heller, P. L. and Wing, Scott L. 2006. History and causes of post-Laramide relief in the Rocky Mountain orogenic plateau, Geological Society of America Bulletin, 118(3):393-405

Wing, Scott L. 2005. "A paleontological perspective on plant extinction". Pp. 43-53 in Plant Conservation A Natural History Approach (Krupnick, Gary A. and Kress, W. John). University of Chicago Press

Wing, Scott L. 2005. Transient floral change and rapid global warming at the Paleocene-Eocene boundary, Science, 310(5750):993-996

Greenwood, D. R., Wilf, Peter, Wing, Scott L. and Christophel, D. C. 2004. Paleotemperature Estimation Using Leaf-Margin Analysis: Is Australia Different?, Palaios, 19:129-142

Barnosky, Anthony D., Koch, Paul L., Feranec, Robert S., Wing, Scott L. and Shabel, Alan B. 2004. Assessing the causes of Late Pleistocene extinctions on the continents, Science, 306:70-75

Fricke, H. and Wing, Scott L. 2004. Oxygen isotope and paleobotanical estimates of temperature and A'18O latitudinal gradients over North America during the early Eocene, American Journal of Science, 304:612-635

DiMichele, William A., Behrensmeyer, Anna K., Olszewski, T. D., Labandeira, Conrad C., Pandolfi, John M., Wing, Scott L. and Bobe, Rene. 2004. Long-term stasis in ecological assemblages: evidence from the fossil record, Annual Review of Ecology, Evolution, and Systematics, 35:285-322

Wing, Scott L. 2004. "Mass Extinctions in Plant Evolution". Pp. 61-97 in Extinctions in the History of Life (Taylor, Paul D.). Cambridge University Press

Koch, P. L., Clyde, W. C., Heppel, R. P., Fogel, Marilyn L., Wing, Scott L. and Zachos, J. C. 2003. Isotopic records of carbon cycle and climate change across the Paleocene-Eocene boundary from the Bighorn Basin, Wyoming, Special Papers / Geological Society of America, 369:49-64

Wing, Scott L., Harrington, Guy J., Bowen, G. J. and Koch, P. L. 2003. Floral change during the Initial Eocene Thermal Maximum in the Powder River Basin, Wyoming, Special Papers / Geological Society of America, 369:425-440

Royer, Dana L., Hickey, Leo J. and Wing, Scott L. 2003. Ecological conservatism in the "living fossil" Ginkgo, Paleobiology, 29(1):84-104

Wilf, Peter, Caneo, N. R., Johnson, Kirk R., Hicks, J. F., Wing, Scott L. and Obradovich, J. D. 2003. High plant diversity in Eocene South America: evidence from Patagonia, Science, 300:122-125

Wing, Scott L. and Harrington, G. J. 2001. Floral response to rapid warming in the earliest Eocene and implications for concurrent faunal change, Paleobiology, 27(3):539-563

Royer, D. L., Wing, Scott L., Beerling, D. J., Jolley, D. W., Koch, P. L., Hickey, Leo J. and Berner, R. A. 2001. Paleobotanical evidence for near present-day levels of atmospheric CO2 during part of the tertiary, Science, 292(5525):2310-2313

Wing, Scott L. 2001. Hot times in the Bighorn Basin, Natural History, 110(3):48-54

Huber, Brian T., MacLeod, Kenneth G. and Wing, Scott L. (eds.).2000. Warm Climates in Earth History. Cambridge University Press

Sewall, J. O., Sloan, L. C., Huber, M. and Wing, Scott L. 2000. Climate sensitivity to changes in land surface characteristics, Global and Planetary Change, 26(4):445-465

Wing, Scott L. 2000. Cool, warm, cool, warm - climate oscillation and floral change during the Paleocene, Gff, 122:182-183

Wing, Scott L. 2000. Evolution and Expansion of Flowering Plants, Paleontological Society Papers, 6:209-232

Wilf, Peter, Wing, Scott L., Greenwood, D. R. and Greenwood, C. L. 1999. Using fossil leaves as paleoprecipitation indicators: An Eocene example: Reply, Geology, 27(1):92-92

Manchester, S. R., Dilcher, D. L. and Wing, Scott L. 1998. Attached leaves and fruits of myrtaceous affinity from the middle Eocene of Colorado, Review of Palaeobotany and Palynology, 102(3-4):153-163

Wing, Scott L. and Boucher, L. D. 1998. Ecological aspects of the Cretaceous flowering plant radiation, Annual Review of Earth and Planetary Sciences, 26:379-421

Wilf, Peter, Wing, Scott L., Greenwood, D. R. and Greenwood, C. L. 1998. Using fossil leaves as paleoprecipitation indicators: An Eocene example, Geology, 26(3):203-206

Davies-Vollum, K. S. and Wing, Scott L. 1998. Sedimentological, taphonomic, and climatic aspects of Eocene swamp deposits (Willwood Formation, Bighorn Basin, Wyoming), Palaios, 13(1):28-40

Wing, Scott L. 1998. "Tertiary Vegetation of North America as a Context for Mammalian Evolution". Pp. 37-60 in Evolution of Tertiary Mammals of North America, Volume 1: Terrestrial Carnivores, Ungulates, and Ungulatelike Mammals (Janis, Christine M., Scott, Kathleen M. and Jacobs, Louis L.). Cambridge University Press

Wing, Scott L. 1997. "Global Warming and Plant Species Richness: a Case Study of the Paleocene/Eocene Boundary". Pp. 163-185 in Biodiversity II : Understanding and Protecting our Biological Resources (Reaka-Kudla, Marjorie L., Wilson, Don E. and Wilson, Edward O.). Joseph Henry Press

Damuth, J., Behrensmeyer, Anna K., DiMichele, William A., Labandeira, Conrad C., Potts, Richard and Wing, Scott L.1997. ETE Database Manual. Evolution of Terrestrial Ecosystems Consortium.

Wing, Scott L. 1996. The Republic Highlands, Washington Geology, 24(2):40

Wing, Scott L. 1996. "Global Warming and Plane Species Richness—a Case Study of the Paleocene/Eocene Boundary". Pp. 163-185 in Biodiversity II: Understanding and Protecting our Natural Resources (Wilson, E. O., Wilson, Don E. and Reaka-Kudla, M. L.). National Academy Press

Wing, Scott L. and Greenwood, D. 1996. Eocene continental climates and latitudinal temperature gradients: Reply, Geology, 24(11):1054-1055

Greenwood, D. R. and Wing, Scott L. 1995. Eocene Continental Climates and Latitudinal Temperature-Gradients, Geology, 23(11):1044-1048

Wing, Scott L., Alroy, J. and Hickey, Leo J. 1995. Plant and Mammal Diversity in the Paleocene to Early Eocene of the Bighorn Basin, Palaeogeography, palaeoclimatology, palaeoecology, 115(1-4):117-155

Wing, Scott L. and DiMichele, William A. 1995. Conflict between local and global changes in plant diversity through geological time, Palaios, 10:551-564

Greenwood, D. R., Wing, Scott L. and Behrensmeyer, Anna K. 1993. Determining Paleoclimates, Science, 260(5106):278-279

Wing, Scott L. and Wolfe, J. A. 1993. Stable-Isotope Study of Fluid Inclusions in Fluorite from Idaho - Implications for Continental Climates during the Eocene - Comment, Geology, 21(11):1051-1051

Wing, Scott L. and Greenwood, D. R. 1993. Fossils and Fossil Climate - the Case for Equable Continental Interiors in the Eocene, Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences, 341(1297):243-252

Wing, Scott L., Hickey, Leo J. and Swisher, C. C. 1993. Implications of an Exceptional Fossil Flora for Late Cretaceous Vegetation, Nature, 363(6427):342-344

Wing, Scott L. 1992. High-Resolution Leaf X-Radiography in Systematics and Paleobotany, American Journal of Botany, 79(11):1320-1324

Burnham, Robyn, Wing, Scott L. and Parker, Geoffrey G. 1992. Reflection of deciduous forest communities in leaf litter: Implications for autochthonous litter assemblages from the fossil record, Paleobiology, 18(1):30-49

Wing, Scott L. and Sues, Hans-Dieter. 1992. "Mesozoic and Early Cenozoic Terrestrial Ecosystems". Pp. 327-416 in Terrestrial Ecosystems Through Time : Evolutionary Paleoecology of Terrestrial Plants and Animals (Behrensmeyer, Anna K., Damuth, John D., DiMichele, William A., Potts, Richard, Sues, Hans-Dieter and Wing, Scott L.). University of Chicago Press

Wing, Scott L., Bown, T. M. and Obradovich, J. D. 1991. Early Eocene Biotic and Climatic-Change in Interior Western North-America, Geology, 19(12):1189-1192

Archibald, J. D. and Wing, Scott L. 1991. Equable Climates during Earth History - Comment, Geology, 19(5):539-540

McCartan, L., Tiffney, B. H., Wolfe, J. A., Ager, Thomas A., Wing, Scott L., Sirkin, L. A., Ward, L. W. and Brooks, J. 1990. Late Tertiary Floral Assemblage from Upland Gravel Deposits of the Southern Maryland Coastal-Plain, Geology, 18(4):311-314

Wing, Scott L. 1987. Eocene and Oligocene Floras and Vegetation of the Rocky Mountains, Annals of the Missouri Botanical Garden, 74(4):748-784

Wing, Scott L. and Tiffney, B. H. 1987. The Reciprocal Interaction of Angiosperm Evolution and Tetrapod Herbivory, Review of Palaeobotany and Palynology, 50(1-2):179-210

Wing, Scott L. 1987. "Interactions of Angiosperms and Herbivorous Tetrapods through Time". Pp. 203-224 in The Origins of Angiosperms and their Biological Consequences (Friis, E. M., Chaloner, W. G. and Crane, P. R.). Cambridge University Press

Brunet, M., Coppens, Y., Pilbeam, D., Djallo, S., Behrensmeyer, Anna K., Brillanceau, A., Downs, W., Duperon, M., Ekodeck, G., Flynn, L., Heintz, E., Hell, J., Jehenne, Y., Martin, L., Mosser, C., Salardcheboldaeff, M., Wenz, S. and Wing, Scott L. 1986. Cenozoic and Cretaceous Fresh-Water Deposits in Cameroon - Preliminary-Results of a Paleontological Prospection, Comptes Rendus De L Academie Des Sciences Serie Ii, 303(5):425-428

Wing, Scott L. and Bown, T. M. 1985. "Fine Scale Reconstruction of the Late Paleocene-Early Eocene Paleogeography in the Bighorn Basin of Northern Wyoming". Pp. 93-105 in Cenozoic Paleogeography of the West Central United States, Rocky Mountain Symposium (Flores, R. M. and Kaplan, S.). Rocky Mountain Section, Society of Economic Paleontologists and Mineralogists

Wing, Scott L. 1984. Impressions of Fallen Leaves, Earth Science, 37(4):16-17

Wing, Scott L. 1984. Review of "Catastrophes and Earth History", Earth Sciences History, 3(2):190-191

Wing, Scott L. 1984. A New Basis for Recognizing the Paleocene Eocene Boundary in Western Interior North-America, Science, 226(4673):439-441

Wing, Scott L. and Hickey, Leo J. 1984. The Platycarya Perplex and the Evolution of the Juglandaceae, American Journal of Botany, 71(3):388-411

Wing, Scott L. 1984. Relation of Paleovegetation to Geometry and Cyclicity of some Fluvial Carbonaceous Deposits, Journal of Sedimentary Petrology, 54(1):52-66